I am currently a Professor of Neurobiology at Harvard Medical School and was Director of the Harvard PhD Program in Neuroscience from 2009 to 2014. I was raised in the Upper Peninsula of Michigan and attended DePauw University where I received a B.A. in Chemistry in 1983. I attended Harvard Medical School (HMS), where I discovered the joys of visual neurophysiology working with Professors David Hubel and Margaret Livingstone. After receiving the M.D. degree in 1988, I continued as a postdoctoral fellow in the Hubel/Livingstone lab, undertook a second postdoc with William Newsome at Stanford and then returned to HMS in 1995 as an Assistant Professor in the Department of Neurobiology. My laboratory studies cortical visual processing in nonhuman primates, with a particular interest in the nature of cortico-cortical feedback. I have a daughter just out of college, and I live in Jamaica Plain with my wife and two very active dogs. My hobbies include bicycling, ocean swimming and going to the opera.